Analyses on Arbor Tree Species Distribution Pattern and Stand Spatial Structure

doi:10.13466/j.cnki.lyzygl.2020.01.008

Abstract

Abstract:

Trees are an important factor in the stablity of forest ecosystem.The analysis of stand structure is of great significance to the stability protection and sustainable management of forest ecosystem.In this paper,the ninth national forest resources inventory data and special survey data of vegetation quadrat were used for cluster analysis to calculate tree species distribution pattern index and stand structure characteristic index,and the Gaussian distribution was used to fit the probability distribution of structure characteristic index,so as to provide a reference for forest ecosystem protection and management.The results show that the distribution of main tree species is clustered.The dispersion degree of arbor forest mixture index Mc_i is greater than that of tree species spatial structure TSS index. TSS index distribution shows the left truncated normal distribution,and the Mc_i index distribution shows the inverted "J" distribution.The fluctuation and continuous stability of the index may be related to artificial afforestation and natural forest protection.

Key words: distribution of tree species, spatial structure, mixed degree, Gaussian distribution, forest ecology

CLC Number:

S718.5

Heng WU, Liyan ZHU, Hailiang WANG, Zhijun LIU. Analyses on Arbor Tree Species Distribution Pattern and Stand Spatial Structure[J]. FOREST RESOURCES WANAGEMENT, 2020, (1): 54-61.

Figures/Tables 5

Tab.1

Descriptive statistics of the calculated results of sample TSS and Mci indicators

样本	样地 TSS 指标				样地 Mc_i 指标
样本	平均值	标准差	最小值	最大值	平均值	标准差	最小值	最大值
总体	0.3724	53.6110	0.0008	1.0000	0.2941	55.9301	0.0005	1.1500
S1	0.2467	6.1558	0.0027	1.0000	0.2167	6.4625	0.0025	1.1500
S2	0.2294	2.6351	0.0094	0.8400	0.1961	2.8435	0.0041	0.9240
S3	0.3461	8.8742	0.0046	0.9558	0.2169	8.7061	0.0007	1.0971
S4	0.2533	6.3763	0.0042	1.0000	0.2204	6.8419	0.0028	1.1500
S5	0.2546	5.5377	0.0068	0.8400	0.2152	5.5688	0.0029	0.9240
S6	0.3046	6.3500	0.0049	0.8400	0.2838	6.8703	0.0008	0.9240
S7	0.4830	13.7076	0.0067	0.9913	0.4573	15.9040	0.0013	1.1323
S8	0.3062	4.8142	0.0061	0.8857	0.2313	4.8408	0.0010	0.9414
S9	0.2603	4.8449	0.0026	0.8667	0.2315	5.2981	0.0029	0.9892
S10	0.3922	8.9108	0.0048	1.0000	0.2547	9.2331	0.0008	1.1500
S11	0.3658	8.7095	0.0040	0.9429	0.3427	9.8094	0.0010	1.0442
S12	0.4582	14.6923	0.0046	1.0000	0.3041	15.9005	0.0007	1.1500
S13	0.4718	14.2195	0.0045	1.0000	0.3154	15.6600	0.0007	1.1500
S14	0.3321	6.9700	0.0031	0.8400	0.3079	7.5525	0.0008	0.9735
S15	0.2432	5.9844	0.0026	1.0000	0.2096	6.5586	0.0023	1.1294
S16	0.3713	11.8896	0.0047	1.0000	0.2588	12.2596	0.0007	1.1500
S17	0.4994	11.3967	0.0055	1.0000	0.3632	13.3911	0.0009	1.1500
S18	0.4552	13.8417	0.0048	0.9613	0.3304	15.4725	0.0008	1.1193
S19	0.4688	8.9414	0.0043	0.9788	0.3208	9.8779	0.0007	1.0738
S20	0.4496	11.7760	0.0032	0.9733	0.3382	13.0862	0.0005	1.0905
S21	0.3390	5.3460	0.0115	0.9377	0.2095	5.0988	0.0018	0.8590
S22	0.4638	6.1121	0.0104	0.8750	0.4537	7.0285	0.0067	0.9683
S23	0.2789	8.9589	0.0058	1.0000	0.2473	9.6130	0.0020	1.1317
S24	0.4433	11.3525	0.0045	0.9653	0.3215	12.5864	0.0007	1.1012
S25	0.2705	9.4448	0.0029	0.9489	0.2367	9.7900	0.0012	0.8575
S26	0.5377	6.9152	0.0078	0.9621	0.1930	5.9244	0.0013	1.0555
S27	0.3360	8.1203	0.0053	0.8400	0.3094	8.9138	0.0023	0.9240
S28	0.3005	7.7390	0.0016	0.8857	0.2688	8.2757	0.0011	0.9342
S29	0.1883	2.7151	0.0075	0.6515	0.1460	2.6359	0.0012	0.6518
S30	0.3572	4.9478	0.0106	0.9765	0.2255	4.9795	0.0017	0.9936
S31	0.2078	4.1634	0.0008	0.8400	0.1703	4.1670	0.0011	0.9240

Tab.1

Fig.1

Tab.2

Tab.3

Fig.2

References 25

[1]	国家林业和草原局. 中国森林资源报告(2014-2018)[M]. 北京: 中国林业出版社, 2019, 48-76.
[2]	赵中华, 惠刚盈, 胡艳波 , 等. 树种多样性计算方法的比较[J]. 林业科学, 2012,48(11):1-8. doi: 10.11707/j.1001-7488.20121101
[3]	惠刚盈, 胡艳波, 赵中华 . 结构化森林经营研究进展[J]. 林业科学研究, 2018,31(1):85-93.
[4]	Hui Gangying, Zhao Xiuhai, Zhao Zhonghua , et al. Evaluating tree species spatial diversity based on neighborhood relationships[J]. Forest Science, 2011, volume 57(4):292-300.
[5]	赵中华, 惠刚盈, 刘文桢 , 等. 小陇山林区2种锐齿栎次生林林分的结构特征[J]. 西北农林科技大学学报:自然科学版, 2019,47(8):75-82.
[6]	LexerD N L, Eid T . An evaluation of different diameter diversity indices based on criteria related to forest management planning[J]. Forest Ecology and Management, 2006,222(1-3):17-28. doi: 10.1016/j.foreco.2005.10.046
[7]	汤孟平, 娄明华, 陈永刚 , 等. 不同混交度指数的比较分析[J]. 林业科学, 2012,48(8):46-53. doi: 10.11707/j.1001-7488.20120808
[8]	Onaindia M, Dominguez I, Albizu I , et al. Vegetation diversity and vertical structure as indicators of forest disturbance[J]. Forest Ecology and Management, 2004,195(3):341-354. doi: 10.1016/j.foreco.2004.02.059
[9]	Shimatani K . Multivariate point processes and spatial variation of species diversity[J]. Forest Ecology and Management, 2001,142(1-3):215-229. doi: 10.1016/S0378-1127(00)00352-2
[10]	徐炜, 马志远, 井新 , 等. 生物多样性与生态系统多功能性:进展与展望[J]. 生物多样性, 2016,24(1):55-71. doi: 10.17520/biods.2015091
[11]	张景慧, 黄永梅 . 生物多样性与稳定性机制研究进展[J]. 生态学报, 2016,36(13):3859-3870. doi: 10.5846/stxb201411062196
[12]	安慧君, 张智杰, 常峥 , 等. 红花尔基林业局森林生态功能评价[J]. 林业资源管理, 2016(5):131-137.
[13]	汤孟平, 唐守正, 李希菲 , 等. 树种组成指数及其应用[J]. 林业资源管理, 2003(2):33-36.
[14]	雷相东, 唐守正 . 林分结构多样性指标研究综述[J]. 林业科学, 2002(3):140-146.
[15]	姚爱静, 朱清科, 张宇清 , 等. 林分结构研究现状与展望[J]. 林业调查规划, 2005(2):70-76.
[16]	汤孟平, 唐守正, 雷相东 , 等. 两种混交度的比较分析[J]. 林业资源管理, 2004(4):25-27.
[17]	于亦彤, 王新杰, 刘丽 , 等. 基于Voronoi图的不同择伐强度下云冷杉林结构分析[J]. 西北农林科技大学学报:自然科学版, 2019,47(9):69-78.
[18]	龚直文, 亢新刚, 顾丽 , 等. 天然林林分结构研究方法综述[J]. 浙江林学院学报, 2009,26(3):434-443.
[19]	Pommerening A . Evaluating structural indices by reversing forest structural analysis[J]. Forest Ecology and Management, 2006,224(3):266-277. doi: 10.1016/j.foreco.2005.12.039
[20]	Neumann M, Starlinger F . The significance of different indices for stand structure and diversity in forests[J]. Forest Ecology and Management, 2001,145(1-2):91-106. doi: 10.1016/S0378-1127(00)00577-6
[21]	Gadow K v, Zhang Chunyu, Wehenkel C , et al. Forest structure and diversity[M]. Continuous Cover Forestry:Springer Netherlands, 2012: 29-83.
[22]	吴建强, 王懿祥, 杨一 , 等. 干扰树间伐对杉木人工林林分生长和林分结构的影响[J]. 应用生态学报, 2015,26(2):340-348.
[23]	李婷婷, 陆元昌, 姜俊 , 等. 马尾松人工林森林经营模式评价[J]. 西北林学院学报, 2015,30(1):164-171.
[24]	惠刚盈, Klaus von, Gadow,等. 结构化森林经营原理[M]. 北京: 中国林业出版社, 2016.
[25]	Graz F P . The behaviour of the measure of surround in relation to the diameter and spatial structure of a forest stand[J]. European Journal of Forest Research, 2008,127(2):165-171. doi: 10.1007/s10342-007-0193-3

树种	扩散系数 C	聚集度指标 I	CassieRM指标 C_A	平均拥挤度与平均密度的比值 R
华山松 Pinus armandii	80.27	79.27	4.52	5.52
杉木 Cunninghamia lanceolata	74.55	73.55	1.87	2.87
柳杉 Cryptomeria fortunei	74.41	73.41	3.24	4.24
桉树 Eucalyptus robusta	63.43	62.43	1.29	2.29
油松 Pinus tabuliformis	54.74	53.74	1.90	2.90
栎类 Quercus acutissima	52.07	51.07	1.91	2.91
云南松 Pinus yunnanensis	51.17	50.17	1.34	2.34
马尾松 Pinus massoniana Lamb.	48.93	47.93	1.99	2.99
栓皮栎 Quercus variabilis	47.67	46.67	2.40	3.40
柏木 Cupressus funebris	46.34	45.34	2.33	3.33
木荷 Schima superba	40.30	39.30	2.78	3.78
落叶松 Larix gmelinii(Rupr.) Kuzen.	39.29	38.29	1.37	2.37
刺槐 Robinia pseudoacacia	37.57	36.57	3.02	4.02
湿地松 Pinus elliottii	36.82	35.82	1.42	2.42
桦木 Betula	34.74	33.74	3.14	4.14
白桦 Betula platyphylla Suk.	33.47	32.47	1.68	2.68
杨树 Populus L.	31.78	30.78	1.61	2.61
云杉 Picea asperata Mast.	31.41	30.41	1.19	2.19
樟木 Cinnamomum camphora	31.24	30.24	4.53	5.53
冷杉 Abies fabri(Mast.) Craib	27.59	26.59	1.47	2.47
柳树 Salix	23.25	22.25	3.25	4.25
高山松 Pinus densata	22.29	21.29	0.64	1.64
楠木 Phoebe zhennan	20.31	19.31	2.37	3.37
榆树 Ulmus pumila L.	18.89	17.89	3.04	4.04
枫香 Liquidambar formosana	13.94	12.94	2.18	3.18

指数	参数	拟合值	标准估计误差	t 值	p 值
TSS	a₁	0.0337	0.0045	7.4826	<0.0001
	a₂	0.2941	0.0474	6.2082	<0.0001
	a₃	0.2930	0.0175	16.7788	<0.0001
	a₄	-0.0006	0.0049	-0.1215	0.9039
Mc_i	a₁	0.0469	0.0241	1.9458	0.0582
	a₂	0.5112	0.3662	1.3962	0.1698
	a₃	-0.0937	0.3697	-0.2536	0.8011
	a₄	-0.0047	0.0162	-0.2880	0.7748